As the only member of the pop music club not to graduate, the responsibility of recruiting new members has fallen to Azusa. Ui and Jun have agreed to help out their friend, but they might be more interested in having actual afterschool teatime than making music. Even when Azusa secures the necessary four members to keep the club alive, the lack of musical experience (and musical instruments) is going to be a challenge! Will the high school pop music club survive to take the stage once more?!

At the old-fashioned Japanese boarding house Narutakisou, Shiratori lives with a diverse yet strangely-similar group of tenants. Under the tender care of landlady Aoba Kozue, Narutakisou provides a home to people whose personal lives are all painful in one way or another; yet together, somehow, they make up a true family. Not only does Shiratori find a home at Narutakisou, but he also finds himself in love with a girl who is far more than the sum of her parts.

Mushi are another form of life which live parallel to our own natural world. The source of many myths and superstitions, they exist according to their own natural laws, and can be both beneficial and dangerous depending on circumstances.  Ginko, a traveling mushishi, investigates and helps out wherever the mushi have become a problem. Yet Ginko himself is marked by the creatures he researches, and he carries the weight of his past with him.